We studied soil hydraulic conductivity (K) and porosity in five combinations of soil tillage and cover crop management systems. Treatments were winter wheat (Triticum aestivum L.) grown on a conventionally tilled soil (CT), on a no-till soil (NT), and on an NT with three different cover crops: red fescue (Festuca rubra L.; Fr), bird's-foot-trefoil (Lotus corniculatus L.; Lc) and alfalfa (Medicago sativa L.; Ms). Measurements were made on a loamy soil in Grignon, France, in November 2004, May 2005 and October 2005. K and mean size of hydraulically active pores were measured in situ at three water potentials (22120.6, 22120.2 and 22120.05 kPa) at the soil surface and at 10 cm depth. In November 2004 and May 2005, pore space was described using 2D image analysis of pores on undisturbed soil samples in the 0201310 cm layer and in the 10201320 cm layer. The major differences were caused by soil tillage that created two heterogeneous soil layers and increased K in the 0201310Â cm layer relative to NT. The effects of cover crop on K and porosity were not affected by the root type: there were no major differences between the grass cover crop (fibrous-root type) and the leguminous ones (tap-root type). However, we recorded larger functional pores and more tubules in the no-till treatments with a cover crop, compared with the no-till treatment without cover crop; this was probably the result of root activity. Although these changes generally did not result in larger values of K, they participated in the maintenance of soil structure and K over time.