Net ecosystem productivity (NEP) was measured on shortgrass steppe (SGS) vegetation at the USDA Central Plains Experimental Range in northeastern Colorado from 2001 to 2003. Large year-to-year differences were observed in annual NEP, with >95% of the net carbon uptake occurring during May and June. Low precipitation during the 2002 April to June time period greatly reduced annual net carbon uptake. Large precipitation events (>10 mm day -1) promoted carbon uptake, while small precipitation events (5 mm having a similar increase in respiration (>3.00 g m Cm -2 day -1). In addition, the size of the heterotrophic respiration pulse is independent of both the amount of time since the last rainfall event and the time of occurrence during the growing season.